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Species code:	IGNH4
Taxonomy:	Archaea; Crenarchaeota; Thermoprotei; Desulfurococcales; Desulfurococcaceae; Ignicoccus (TaxID: 453591) [NEWT/ NCBI]
Description:	An anaerobic chemolithoautotrophic and hyperthermophilic archaeon, Ignicoccus hospitalis was isolated from a submarine hydrothermal system off the coast of Iceland. Optimal growth occurs at 90 degrees C utilizing molecular hydrogen, elemental sulfur, and carbon dioxide as the energy substrate, the electron acceptor, and the carbon source, respectively. It is the obligatory host of Nanoarchaeum equitans; up to a dozen N.equitans can be found on the surface of I.hospitalis without doing any apparent harm to the host. Ignicoccus species are unique among the Archaea in having two cell membranes; together with the cytoplasmic membrane, it encloses a huge periplasmic space, in which membrane-bound vesicles are found. I.hospitalis at 1.3 Mb has one of the smallest genomes among free-living bacteria and archaea, and shows signs of gene exchange with N.equitans as well as gene acquisition from Euryarchaeota and bacteria (adapted from PMID 19000309).
Properties:	Presence of flagella: Yes Human pathogen: No Interaction: Archaeal host for Naoarchaeum equitans Number of membranes: 2 Number of inteins:0
Statistics:	Number of IGNH4 entries in the UniProt Knowledgebase: 1434 (124 in UniProtKB/Swiss-Prot + 1310 in UniProtKB/TrEMBL)

Genome structure:	Chromosome EMBL accession number CP000816 (circular; 1,297,538 bp) (download entry)
Reference(s):	[1] PubMed=19000309; [ NCBI , EBI , Israel , Japan ] Podar M., Anderson I., Makarova K.S., Elkins J.G., Ivanova N., Wall M.A., Lykidis A., Mavromatis K., Sun H., Hudson M.E., Chen W., Deciu C., Hutchison D., Eads J.R., Anderson A., Fernandes F., Szeto E., Lapidus A., Kyrpides N.C., Saier M.H. Jr., Richardson P.M., Rachel R., Huber H., Eisen J.A., Koonin E.V., Keller M., Stetter K.O. ; "A genomic analysis of the archaeal system Ignicoccus hospitalis-Nanoarchaeum equitans."; Genome Biol. 9:R158.1-R158.18(2008).
Web links:	Entrez Genome Project: http://www.ncbi.nlm.nih.gov/sites/entrez?Db=genomeprj&cmd=ShowDetailView&TermToSearch=13914 GIB (DDBJ): http://gib.genes.nig.ac.jp/single/index.php?spid=Ihos_KIN4I  EBI Proteome Analysis page

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